Middle Triassic pteridosperms (Pinophyta) of the Timan–Pechora basin

Resumo

The collection of fossil plants sampled by geologists from VNIGRI at the end of the 20th century from Triassic continental sections drilled by many wells and cropping out in several natural localities and stored at the Museum of Petroleum Geology and Paleontology of the same institute was critically revised. The use of the epidermal method for the study of plant remains with consideration of recent publications dedicated to continental sections of Central Europe made it possible to substantially broaden the taxonomic composition of the Triassic flora and first specify the composition of its pteridosperm representatives. Unlike the Triassic floras of Western Europe, the pteridosperms the Pechora region appeared to be relatively diverse. They number 37 species of 11 genera, which are confined to the upper part of the Triassic sequence: Anguran and Naryan-Mar formations and their analogs. The Middle Triassic, mainly, Ladinian, age of these formations is reliably substantiated both by paleontological (vertebrate and palynological) data and by results of the comparative analysis of the Anguran–Naryan-Mar taphofloras and coeval European type floras dated back to the Anisian–Ladinian by marine faunal remains. The stratigraphic significance of pteridospermous plant remains becomes undoubted for continental sections of the Timan–Pechora basin, while the genera Scytophyllum, Kalantarium, and Kirjamkenia may be considered with respect to their diversity and abundance as representing orthostratigraphic taxa.